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Gratiana boliviana
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Egg
Female: notice white oviducts
Male: notice orange testis
Larva frass
Larva feeding
Pupae
Larvae without frass
Left: mature adult;
Right: young adult
Left: mature adult; Right: young adult
Mature adult
Old adult
Left: Mature adult, Right: Old adult
Adults mating
Larvae feeding
Feeding damage
Feeding damage
Adult feeding
Agent Identification
Host specificity
Impact on TSA
Spread of Gratiana boliviana
Mapping Gratiana releases in the USA
How can I obtain beetles?
Pictures